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The Joker has been referred to as the Clown Prince of Crime, the Harlequin of Hate, and the Ace of Knaves. Throughout the evolution of the DC universe, interpretations and incarnations of the Joker have taken two forms. The original and currently dominant image is of a fiendishly intelligent lunatic with a warped, sadistic sense of humor. The other interpretation of the character, popular in the late 1940s through 1960s comic books as well as the 1960s television series, is that of an eccentric but harmless prankster and thief. The 1990s cartoon Batman: The Animated Series blended these two aspects, although most interpretations tend to embrace one characterization or the other.[37]

The Joker's victims have included men, women, children, and even his own henchmen. In the graphic novel The Joker: Devil's Advocate, the Joker is reported to have killed well over 2,000 people. Despite having murdered enough people to get the death penalty thousands of times over, he is always found not guilty by reason of insanity.[39] In the Batman story line "War Crimes", this continued ruling of insanity is in fact made possible by the Joker's own dream team of lawyers. He is then placed in Arkham Asylum, from which he appears able to escape at will, going so far as to claim that it's just a resting ground in between his "performances".

There have been times when Batman has been tempted to put the Joker down once and for all, but has relented at the last minute. After capturing the Joker in one story, he threatens to kill his old foe, but then says, "But that would give you the final victory, making me into a killer like yourself!" Conversely, the Joker has given up many chances to kill Batman. The Joker's obsession with Batman is unique compared to other villains, in that he is portrayed as enjoying their battles and constantly mocking him, hinting he may want to make Batman as insane as he is.

The Joker is renowned as Batman's greatest enemy.[40] While other villains rely on tried-and-true methods to commit crimes (such as Mr. Freeze's freeze gun or Poison Ivy's toxic plants), Joker has a variety of weapons at his disposal. For example, the flower he wears in his lapel sprays (at any given time) acid, poisonous laughing gas, or nothing at all. In Batman Beyond: Return of the Joker and much earlier in "Dreadful Birthday, Dear Joker!" (Batman #321), the Joker has a gun which at first shoots a flag saying "BANG!", but then, with another pull of the trigger, the flag fires and impales its target.[35][41] His most recurring gadget is a high-voltage hand-buzzer, which he uses to electrocute his victims with a handshake. Sometimes he commits crimes just for the fun of it, while on other occasions, it is part of a grand scheme; Batman has been noted to say that the Joker's plans "make sense to him alone." His capricious nature, coupled with his violent streak, makes him the one villain that the DC Universe's other super-villains fear; in the Villains United and Infinite Crisis mini-series, the members of the villains' Secret Society refuse to induct the Joker for this reason. In the mini-series Underworld Unleashed, the Trickster remarks, "When super-villains want to scare each other, they tell Joker stories".

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